Cucumber-Peanut Intercropping: The Effects on Vegetative Growth and Yield Productivity
DOI:
10.29303/jppipa.v12i1.12758Published:
2026-01-25Downloads
Abstract
Intercropping is acknowledged as an effective agricultural intensification strategy for limited land. Planting time is an important factor in intercropping systems because it may intensify canopy overlap and shading, particularly because both crops rely on the C3 photosynthetic pathway, potentially reducing growth and yield. This study examined the effect of different planting times for cucumber (Cucumis sativus L.) and peanut (Arachis hypogaea L.). The study was conducted from October 2023 to February 2024 with a one-factor randomized block design. Five treatments were applied: simultaneous planting and cucumber planting at 1, 2, 3, and 4 weeks after peanuts, each replicated five times. Several data were analyzed using analysis of variance (F-test), followed by Tukey’s HSD test at the 5% significance level. This study confirms that differences in planting time between cucumbers and peanuts significantly affect soil macronutrient dynamics, growth and production, and land use efficiency. Soil macronutrient analysis showed that simultaneous planting (P1) resulted the highest soil nitrogen increase (0.38%). However, this treatment also caused a reduction in phosphorus to 15.15 mg per 100 g of soil and potassium to 13.48 mg per 100 g of soil. In contrast, excessive delay (P5) caused nitrogen to drop to 0.11% and phosphorus to decline to 14.19 mg per 100 g soil, despite potassium remaining relatively high (17.87 mg per 100 g of soil). Simultaneous planting produced the highest vegetative growth of cucumber, with an average of 40.68 leaves per plant, the greatest fruit length (22.16 cm), individual fruit weight (414.55 g), and total fruit weight per plant (2498.39 g). Delaying cucumber planting resulted in the lowest value of 828.45 g in P5. The highest pod weight was recorded in P1 at 62.46 g per plant. In contrast, planting cucumbers one week after peanuts (P2) resulted in the lowest peanut yield, at only 47.14 g per plant. All treatments achieved LER values greater than 1, confirming the superiority of intercropping over monoculture. The highest LER was obtained in P1 (1.61). In conclusion, simultaneous planting of cucumbers and peanuts provides the most optimal balance between soil nutrient utilization, crop growth, yield, and land use efficiency. Therefore, synchronized planting time is a key management strategy for improving productivity and sustainability in cucumber–peanut intercropping systems under limited land conditions.
Keywords:
Intercropping system Land-use efficiency Monoculture system Planting timeReferences
Ad Adhikari, P., Araya, H., Aruna, G., Balamatti, A., Banerjee, S., Baskaran, P., Barah, B. C., Behera, D., Berhe, T., Boruah, P., Dhar, S., Edwards, S., Fulford, M., Gujja, B., Ibrahim, H., Kabir, H., Kassam, A., Khadka, R. B., Koma, Y. S., Natarajan, U. S., Perez, R., Sen, D., Sharif, A., Singh, G., Styger, E., Thakur, A. K., Tiwari, A., Uphoff, N., & Verma, A. (2018). System of Crop Intensification for More Productive, Resource-conserving, Climate-resilient, and Sustainable Agriculture: Experience with Diverse Crops in Varying Agroecologies. International Journal of Agricultural Sustainability, 16(1), 1–28. https://doi.org/10.1080/14735903.2017.1402504 DOI: https://doi.org/10.1080/14735903.2017.1402504
Aksarah, A., Noer, H., & Mitrayani, D. (2022). Pengaruh Waktu Tanam terhadap Pertumbuhan dan Hasil Tanaman Kacang Tanah yang Ditumpangsarikan dengan Tanaman Jagung Manis. Jurnal Agrotech, 12(1), 38–43. https://doi.org/10.31970/agrotech.v12i1.90 DOI: https://doi.org/10.31970/agrotech.v12i1.90
Altieri, M. A., Nicholls, C. I., & Montalba, R. (2017). Technological Approaches to Sustainable Agriculture at A Crossroads: An Agroecological Perspective. Sustainability, 9(3), 349. https://doi.org/10.3390/su9030349 DOI: https://doi.org/10.3390/su9030349
Anjum, S. A., Ashraf, U., Zohaib, A., Tanveer, M., Naeem, M., Ali, I., Tabassum, T., & Nazir, U. (2017). Growth and Development Responses of Crop Plants Under Drought Stress: A Review. Zemdirbyste-Agriculture, 104(3), 267–276. https://doi.org/10.13080/z-a.2017.104.034 DOI: https://doi.org/10.13080/z-a.2017.104.034
Capitanescu, F., Marvuglia, A., Gutiérrez, T. N., & Benetto, E. (2017). Multi-stage Farm Management Optimization Under Environmental and Crop Rotation Constraints. Journal of Cleaner Production, 147, 197–205. https://doi.org/10.1016/j.jclepro.2017.01.076 DOI: https://doi.org/10.1016/j.jclepro.2017.01.076
Clayton, J., Lemanski, K., Solbach, M. D., Temperton, V. M., & Bonkowski, M. (2024). Two-way NxP Fertilisation Experiment on Barley (Hordeum vulgare) Reveals Shift from Additive to Synergistic NP Interactions at critical Phosphorus Fertilisation Level. Frontiers in Plant Science, 15, 1346729. https://doi.org/10.3389/fpls.2024.1346729 DOI: https://doi.org/10.3389/fpls.2024.1346729
Dolezal, J., Fibich, P., Altman, J., Leps, J., Uemura, S., Takahashi, K., & Hara, T. (2020). Determinants of Ecosystem Stability in A Diverse Temperate Forest. Oikos, 129(11), 1692–1703. https://doi.org/10.1111/oik.07379 DOI: https://doi.org/10.1111/oik.07379
Dong, N., Tang, M.-M., Zhang, W.-P., Bao, X.-G., Wang, Y., Christie, P., & Li, L. (2018). Temporal Differentiation of Crop Growth as One of The Drivers of Intercropping Yield Advantage. Scientific Reports, 8(1), 3110. https://doi.org/10.1038/s41598-018-21414-w DOI: https://doi.org/10.1038/s41598-018-21414-w
Ebbisa, A. (2022). Mechanisms Underlying Cereal/Legume Intercropping as Nature-based Biofortification: A Review. Food Production, Processing and Nutrition, 4(1), 19. https://doi.org/10.1186/s43014-022-00096-y DOI: https://doi.org/10.1186/s43014-022-00096-y
Ferioun, M., Zouitane, I., Bouhraoua, S., Elouattassi, Y., Belahcen, D., Errabbani, A., Louahlia, S., Sayyed, R., & El Ghachtouli, N. (2025). Applying Microbial Biostimulants and Drought-tolerant Genotypes to Enhance Barley Growth and yield Under Drought Stress. Frontiers in Plant Science, 15, 1494987. https://doi.org/10.3389/fpls.2024.1494987 DOI: https://doi.org/10.3389/fpls.2024.1494987
Gebru, H. (2015). A Review on The Comparative Advantages of Intercropping to Mono-cropping System. Journal of Biology, Agriculture and Healthcare, 5(9), 1–13. Retrieved from https://iiste.org/Journals/index.php/JBAH/article/view/22307
Gomez, K. A., & Gomez, A. A. (1984). Statistical Procedures for Agricultural Research. John wiley & sons.
Griffiths, M., & York, L. M. (2020). Targeting Root Ion Uptake Kinetics to Increase Plant Productivity and Nutrient Use Efficiency. Plant Physiology, 182(4), 1854–1868. https://doi.org/10.1104/pp.19.01496 DOI: https://doi.org/10.1104/pp.19.01496
Kumar, A., van Duijnen, R., Delory, B. M., Reichel, R., Brüggemann, N., & Temperton, V. M. (2020). Barley Shoot Biomass Responds Strongly to N: P Stoichiometry and Intraspecific Competition, whereas Roots only Alter their Foraging. Plant and Soil, 453(1), 515–528. https://doi.org/10.1007/s11104-020-04626-w DOI: https://doi.org/10.1007/s11104-020-04626-w
Li, C., Hoffland, E., Kuyper, T. W., Yu, Y., Li, H., Zhang, C., Zhang, F., & van der Werf, W. (2020). Yield Gain, Complementarity and Competitive Dominance in Intercropping in China: A Meta-analysis of Drivers of Yield Gain using Additive Partitioning. European Journal of Agronomy, 113, 125987. https://doi.org/10.1016/j.eja.2019.125987 DOI: https://doi.org/10.1016/j.eja.2019.125987
Li, L., Duan, R., Li, R., Zou, Y., Liu, J., Chen, F., & Xing, G. (2022). Impacts of Corn Intercropping with Soybean, Peanut and Millet through Different Planting Patterns on Population Dynamics and Community Diversity of Insects Under Fertilizer Reduction. Frontiers in Plant Science, 13, 936039. https://doi.org/10.3389/fpls.2022.936039 DOI: https://doi.org/10.3389/fpls.2022.936039
Lithourgidis, A. S., Dordas, C. A., Damalas, C. A., & Vlachostergios, D. (2011). Annual Intercrops: An Alternative Pathway for Sustainable Agriculture. Australian Journal of Crop Science, 5(4), 396–410. Retrieved from https://www.cabidigitallibrary.org/doi/full/10.5555/20113157046
Lopez, J. R., Winter, J. M., Elliott, J., Ruane, A. C., Porter, C., & Hoogenboom, G. (2017). Integrating Growth Stage Deficit Irrigation into a Process Based Crop Model. Agricultural and Forest Meteorology, 243, 84–92. https://doi.org/10.1016/j.agrformet.2017.05.001 DOI: https://doi.org/10.1016/j.agrformet.2017.05.001
Matheus, R. (2020). Skenario Pengelolaan Sumber Daya Lahan Kering: Menuju Pertanian Berkelanjutan. Deepublish.
Nasar, J., Ahmad, M., Gitari, H., Tang, L., Chen, Y., & Zhou, X.-B. (2024). Maize/soybean Intercropping Increases Nutrient Uptake, Crop Yield and Modifies Soil Physio-chemical Characteristics and Enzymatic Activities in the Subtropical Humid Region Based in Southwest China. BMC Plant Biology, 24(1), 434. https://doi.org/10.1186/s12870-024-05061-0 DOI: https://doi.org/10.1186/s12870-024-05061-0
Neves, N. M., Paula, R. R., Araujo, E. A., Gorsani Rodrigo, G., Abreu, K. de, & Kunz, S. H. (2022). Contribution of Legume and Non-legume Trees to Litter Dynamics and CNP Inputs in A Secondary Seasonally Dry Tropical Forest. iForest-Biogeosciences and Forestry, 15(1), 8. https://doi.org/10.3832/ifor3442-014 DOI: https://doi.org/10.3832/ifor3442-014
O’Brien, M. J., Pugnaire, F. I., Armas, C., Rodríguez‐Echeverría, S., & Schöb, C. (2017). The Shift from Plant–plant Facilitation to Competition under Severe Water Deficit is Spatially Explicit. Ecology and Evolution, 7(7), 2441–2448. https://doi.org/10.1002/ece3.2875 DOI: https://doi.org/10.1002/ece3.2875
Qin, H.-Y., Wang, C.-B., Wang, P., Yang, S.-C., Li, X.-F., & Li, L. (2025). Intercropping Enhances The Reciprocal Facilitation of N and P Uptake Through complementarity. Plant and Soil, 1–16. Retrieved from https://ui.adsabs.harvard.edu/abs/2025PlSoi.tmp..611Q/abstract DOI: https://doi.org/10.1007/s11104-025-07726-7
Raza, A., Asghar, M. A., Ahmad, B., Bin, C., Hussain, M. I., Li, W., Iqbal, T., Yaseen, M., Shafiq, I., Yi, Z., Ahmad, I., Yang, W., & Weiguo, L. (2020). Agro-techniques for Lodging Stress Management in Maize-soybean Intercropping System—A Review. Plants, 9(11), 1592. https://doi.org/10.3390/plants9111592 DOI: https://doi.org/10.3390/plants9111592
Rusu, M., Filip, M., Cara, I. G., Țopa, D., & Jităreanu, G. (2025). Soil Nutrient Dynamics and Farming Sustainability under Different Plum Orchard Management Practices in The Pedoclimatical Conditions of Moldavian Plateau. Agriculture, 15(5), 509. https://doi.org/10.3390/agriculture15050509 DOI: https://doi.org/10.3390/agriculture15050509
Sadras, V. O., Villalobos, F. J., & Fereres, E. (2017). Crop Development and Growth. Principles of Agronomy for Sustainable Agriculture, 141–158. Retrieved from https://quantalab.ias.csic.es/pdf/2016_Book_PrinciplesOfAgronomyForSustain.pdf DOI: https://doi.org/10.1007/978-3-319-46116-8_11
Schut, A. G., & Reymann, W. (2023). Towards a Better Understanding of Soil Nutrient Dynamics and P and K Uptake. Plant and Soil, 492(1), 687–707. https://doi.org/10.1007/s11104-023-06209-x DOI: https://doi.org/10.1007/s11104-023-06209-x
Selim, M. M. (2020). Introduction to The Integrated Nutrient Management Strategies and Their Contribution to Yield and Soil Properties. International Journal of Agronomy, 2020(1), 2821678. https://doi.org/10.1155/2020/2821678 DOI: https://doi.org/10.1155/2020/2821678
Shah, F., & Wu, W. (2019). Soil and Crop Management Strategies to Ensure Higher Crop Productivity Within Sustainable Environments. Sustainability, 11(5), 1485. https://doi.org/10.3390/su11051485 DOI: https://doi.org/10.3390/su11051485
Shahid, M., Shukla, A. K., Bhattacharyya, P., Tripathi, R., Mohanty, S., Kumar, A., Lal, B., Gautam, P., Raja, R., Panda, B. B., Das, B., & Nayak, A. K. (2016). Micronutrients (Fe, Mn, Zn and Cu) Balance under Long-term Application of Fertilizer and Manure in A Tropical Rice-rice System. Journal of Soils and Sediments, 16(3), 737–747. https://doi.org/10.1007/s11368-015-1272-6 DOI: https://doi.org/10.1007/s11368-015-1272-6
Smith, M. E., & Francis, C. A. (1986). Breeding for Multiple Cropping Systems. MacMillan Publishing Company.
Stomph, T.-J., Dordas, C., Baranger, A., de Rijk, J., Dong, B., Evers, J., Gu, C., Li, L., Simon, J., Jensen, E. S., Wang, Q., Wang, Y., Wang, Z., Xu, H., Zhang, C., Zhang, L., Zhang, W.-P., Bedoussac, L., & van der Werf, W. (2020). Designing Intercrops for high Yield, Yield Stability and Efficient Use of Resources: Are there Principles? Advances in Agronomy, 160(1), 1–50. https://doi.org/10.1016/bs.agron.2019.10.002 DOI: https://doi.org/10.1016/bs.agron.2019.10.002
Su, B., Zhao, G., & Dong, C. (2018). Spatiotemporal Variability of Soil Nutrients and The Responses of Growth during Growth Stages of Winter Wheat In Northern China. PloS One, 13(12), e0203509. https://doi.org/10.1371/journal.pone.0203509 DOI: https://doi.org/10.1371/journal.pone.0203509
Sullivan, P. (1998). Intercropping Principles and Production Practices. ATTRA.
Tang, L., Zhou, L., Su, L., Zhao, H., Zhao, T., & Zheng, Y. (2024). Maize/soybean Intercropping Improve Yield Stability in Red Soil under Different Phosphate Application Rates in Southwest China. Research Square. https://doi.org/10.21203/rs.3.rs-4003136/v1 DOI: https://doi.org/10.21203/rs.3.rs-4003136/v1
Vives-Peris, V., De Ollas, C., Gómez-Cadenas, A., & Pérez-Clemente, R. M. (2020). Root Exudates: From Plant to Rhizosphere and Beyond. Plant Cell Reports, 39(1), 3–17. https://doi.org/10.1007/s00299-019-02447-5 DOI: https://doi.org/10.1007/s00299-019-02447-5
Wang, T., Mu, X., Ni, E., Wang, Q., Li, S., Mao, J., Qing, D., Li, B., Chen, Y., Chen, W., Liang, C., Wu, H., Lu, X., & Tian, J. (2025). Belowground Interaction in Tea/Soybean Intercropping Enhances Tea Quality by Improving Soil Nutrient Dynamics. Plants, 14(11), 1691. https://doi.org/10.3390/plants14111691 DOI: https://doi.org/10.3390/plants14111691
Willey, R. (1979). Intercropping-Its Importance and Research Needs. 1. Competition and Yield Advantages. Field Crop Abstracts, 32(1), 1-10. Retrieved from https://www.scirp.org/reference/referencespapers?referenceid=1465514
Xu, Z., Li, C., Zhang, C., Yu, Y., van der Werf, W., & Zhang, F. (2020a). Intercropping Maize and Soybean Increases Efficiency of Land and Fertilizer Nitrogen Use; A Meta-analysis. Field Crops Research, 246, 107661. https://doi.org/10.1016/j.fcr.2019.107661 DOI: https://doi.org/10.1016/j.fcr.2019.107661
Xue, Y., Xia, H., Christie, P., Zhang, Z., Li, L., & Tang, C. (2016). Crop Acquisition of Phosphorus, Iron and Zinc from Soil in Cereal/Legume Intercropping Systems: A Critical Review. Annals of Botany, 117(3), 363–377. https://doi.org/10.1093/aob/mcv182 DOI: https://doi.org/10.1093/aob/mcv182
Zhang, W.-P., Liu, G.-C., Sun, J.-H., Fornara, D., Zhang, L.-Z., Zhang, F.-F., & Li, L. (2017). Temporal Dynamics of Nutrient Uptake by Neighbouring Plant Species: Evidence from Intercropping. Functional Ecology, 31(2), 469–479. https://doi.org/10.1111/1365-2435.12732 DOI: https://doi.org/10.1111/1365-2435.12732
Zhang, Y., Lei, J., Peng, Y., Chen, X., Li, B., Chen, Y., Xu, Y., Farooq, T. H., Wu, X., Wang, J., & Yan, W. (2024). Impact of Intercropping on Nitrogen and Phosphorus Nutrient Loss in Camellia oleifera Forests on Entisol Soil. Forests, 15(3), 461. https://doi.org/10.3390/f15030461 DOI: https://doi.org/10.3390/f15030461
Zhen, S., van Iersel, M., & Bugbee, B. (2021). Why Far-Red Photons should be Included in The Definition of Photosynthetic Photons and The Measurement of Horticultural Fixture Efficacy. Frontiers in Plant Science, 12, 693445. https://doi.org/10.3389/fpls.2021.693445 DOI: https://doi.org/10.3389/fpls.2021.693445
Zörb, C., Geilfus, C.-M., & Dietz, K.-J. (2019). Salinity and Crop Yield. Plant Biology, 21, 31–38. https://doi.org/10.1111/plb.12884 DOI: https://doi.org/10.1111/plb.12884
License
Copyright (c) 2026 Mardhiana, Eko Hary Pudjiwati, Nurul Chairiyah, Junarius bin Yakobus, Muh. Adiwena

This work is licensed under a Creative Commons Attribution 4.0 International License.
Authors who publish with Jurnal Penelitian Pendidikan IPA, agree to the following terms:
- Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a Creative Commons Attribution 4.0 International License (CC-BY License). This license allows authors to use all articles, data sets, graphics, and appendices in data mining applications, search engines, web sites, blogs, and other platforms by providing an appropriate reference. The journal allows the author(s) to hold the copyright without restrictions and will retain publishing rights without restrictions.
- Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgment of its initial publication in Jurnal Penelitian Pendidikan IPA.
- Authors are permitted and encouraged to post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to productive exchanges, as well as earlier and greater citation of published work (See The Effect of Open Access).






